The amount and rate of production which occur in a given ecosystem over a given time period. It may apply to a single organism, a population, or entire communities and ecosystems. Productivity can be expressed in terms of dry matter produced per area per time (net production), or in terms of energy produced per area per time (gross production = respiration + heat losses + net production). In aquatic systems, productivity is often measured in volume instead of area. Biomass

Ecologists distinguish between primary productivity (by autotrophs) and secondary productivity (by heterotrophs). Plants have the ability to use the energy from sunlight to convert carbon dioxide and water into glucose and oxygen, producing biomass through photosynthesis. Primary productivity of a community is the rate at which biomass is produced per unit area by plants, expressed in either units of energy [joules/(m²)(day)] or dry organic matter [kg/(m²)(year)]. The following definitions are useful in calculating production: Gross primary production (GPP) is the total energy fixed by photosynthesis per unit time. Net primary production (NPP) is the gross production minus losses due to plant respiration per unit time, and it represents the actual new biomass that is available for consumption by heterotrophic organisms. Secondary production is the rate of production of biomass by heterotrophs (animals, microorganisms), which feed on plant products or other heterotrophs. Photosynthesis

Productivity is not spread evenly across the planet. For instance, although oceans cover two-thirds of Earth's surface, they account for only one-third of the Earth's productivity. Furthermore, the factors that limit productivity in the ocean differ from those limiting productivity on land, producing differences in geographic patterns of productivity in the two systems. In terrestrial ecosystems, productivity shows a latitudinal trend, with highest productivity in the tropics and decreasing progressively toward the Poles; but in the ocean there is no latitudinal trend, and the highest values of net primary production are found along coastal regions.